Environmental Change Institute, University of Oxford

Dr Nikolaos Fyllas


Research Groups

Dr Nikolaos Fyllas

Honorary Research Associate


Nikos is a post doctoral researcher in the Ecosystems Group of ECI. His focus is to understand the soil-plant-atmosphere continuum, through the combined use of field observations / measurements and ecosystem models. Nikos mainly works in Tropical and Mediterranean Ecosystems. His work spans from the fields of Functional Ecology and Ecophysiology to that of Ecosystem Modelling and Ecoinformatics.

Nikos is trained as a mechanical engineer (National Technical University of Athens) and holds a PhD in Ecology (Department of Environment, University of the Aegean). He was a Post-doctoral and Marie-Curie fellow in the Ecology and Global Change Cluster, School of Geography, University of Leeds. He has also worked as an independent researcher at the Faculty of Biology, University of Athens.

Current Research

Nikos is developing the Trait-based Forest Simulator (TFS), an individual-based model that uses trait distributions to represent functional tree diversity and alternative plant strategies at simulations of forest dynamics. He works for the GEM-Trait project, linking data from intensively monitored plots with the TFS model in order to explore the geographical patterns of forest productivity across the worlds Tropical regions and the vulnerability of these ecosystems to global change. He also coordinates a forest plot inventory across elevational gradients in Greece, with an ultimate goal to accurately understand their dynamics under global change.



Koufos G, Mavromatis T, Koundouras S, Fyllas NM, Jones GV (2014) Viticulture-climate relationships in Greece: the impacts of recent climate trends on harvest date variation: Viticulture-Climate relationships in Greece. International Journal of Climatology 34:1445-1459.

Kint V, Aertsen W, Fyllas NM, Trabucco A, Janssen E, Özkan K, Muys B (2014) Ecological traits of Mediterranean tree species as a basis for modelling forest dynamics in the Taurus mountains, Turkey. Ecological Modelling 286:53-65.

Christopoulou A, Fyllas NM, Andriopoulos P, Koutsias N, Dimitrakopoulos PG, Arianoutsou M (2014) Post-fire regeneration patterns of Pinus nigra in a recently burned area in Mount Taygetos, Southern Greece: The role of unburned forest patches. Forest Ecology and Management 327:148-156. (Link)

Adamidis GC, Kazakou E, Fyllas NM, Dimitrakopoulos PG (2014) Species Adaptive Strategies and Leaf Economic Relationships across Serpentine and Non-Serpentine Habitats on Lesbos, Eastern Mediterranean. PLOS ONE 9:e96034. (Link)

Fyllas NM, Gloor E, Mercado LM, Sitch S, Quesada CA, Domingues TF, Galbraith DR, Torre-Lezama A, Vilanova E, Ramírez-Angulo H, Higuchi N, Neill DA, Silveira M, Ferreira L, Aymard C. GA, Malhi Y, Phillips OL, Lloyd J (2014) Analysing Amazonian forest productivity using a new individual and trait-based model (TFS v.1). Geoscientific Model Development :1251-1269. doi: 10.5194/gmd-7-1251-2014.(Link)


Atkin OK, Turnbull MH, Zaragoza-Castells J, Fyllas NM, Lloyd J, Meir P, Griffin KL (2013) Light inhibition of leaf respiration as soil fertility declines along a post-glacial chronosequence in New Zealand: an analysis using the Kok method. Plant Soil 367:163-182.

Torello-Raventos M, Feldpausch TR, Veenendaal E, Schrodt F, Saiz G et al (2013) On the delineation of tropical vegetation types with an emphasis on forest/savanna transitions. Plant Ecology & Diversity 6:101-137.


Patiño S, Fyllas NM, Baker TR, Paiva R, Quesada CA, Santos AJB, Schwarz M, ter Steege H, Phillips OL, Lloyd J (2012) Coordination of physiological and structural traits in Amazon forest trees. Biogeosciences 9:775-801.

Swenson NG, Enquist BJ, Pither J, Kerkhoff AJ, Boyle B, Weiser MD, et al (2012) The biogeography and filtering of woody plant functional diversity in North and South America. Global Ecology and Biogeography 21:798-808.

Quesada CA, Phillips OL, Schwarz M, Czimczik CI, Baker TR, Patiño S, Fyllas NM, et al (2012) Basin-wide variations in Amazon forest structure and function are mediated by both soils and climate. Biogeosciences 9:2203-2246.

Fyllas NM, Quesada CA, Lloyd J (2012) Deriving Plant Functional Types for Amazonian forests for use in vegetation dynamics models. Perspectives in Plant Ecology, Evolution and Systematics 14:97-110. (Link)


Mercado LM, Patiño S, Domingues TF, Fyllas NM, Weedon GP, Sitch S, Quesada CA, Phillips OL, Aragão LEOC, Malhi Y, Dolman AJ, Restrepo-Coupe N, Saleska SR, Baker TR, Almeida S, Higuchi N, Lloyd J (2011) Variations in Amazon forest productivity correlated with foliar nutrients and modelled rates of photosynthetic carbon supply. Philosophical Transactions of the Royal Society of London B: Biological Sciences 366:3316-3329.

Quesada CA, Lloyd J, Anderson LO, Fyllas NM, Schwarz M, Czimczik CI (2011) Soils of Amazonia with particular reference to the RAINFOR sites. Biogeosciences 8:1415-1440.

Feldpausch TR, Banin L, Phillips OL, Baker TR, Lewis SL et al (2011) Height-diameter allometry of tropical forest trees. Biogeosciences 8:1081-1106.

Kattge J, Díaz S, Lavorel S, Prentice IC, Leadley P, BÖnisch G, Garnier E, Westoby M, Reich PB, Wright IJ, et al (2011) TRY - a global database of plant traits. Global Change Biology 17:2905-2935.

Papanikolaou AD, Fyllas NM, Mazaris AD, Dimitrakopoulos PG, Kallimanis AS, Pantis JD (2011) Grazing effects on plant functional group diversity in Mediterranean shrublands. Biodiversity and Conservation 20:2831-2843.


Quesada CA, Lloyd J, Schwarz M, Patiño S, Baker TR, Czimczik C, Fyllas NM et al (2010) Variations in chemical and physical properties of Amazon forest soils in relation to their genesis. Biogeosciences 7:1515-1541.

Fyllas NM, Politi PI, Galanidis A, Dimitrakopoulos PG, Arianoutsou M (2010) Simulating regeneration and vegetation dynamics in Mediterranean coniferous forests. Ecological Modelling 221:1494-1504.


Fyllas NM, Troumbis AY, (2009) Simulating vegetation shifts in north-eastern Mediterranean mountain forests under climatic change scenarios. Global Ecology and Biogeography 18, 64-77.

Fyllas NM, Patiño S, Baker TR, Bielefeld Nardoto G, Martinelli LA, Quesada CA, Paiva R, Schwarz M, Horna V, Mercado LM, Santos A, Arroyo L, Jiménez EM, Luizão FJ, Neill DA, Silva N, Prieto A, Rudas A, Silviera M, Vieira ICG, Lopez-Gonzalez G, Malhi Y, Phillips OL, Lloyd J, 2009. Basin-wide variations in foliar properties of Amazonian forest: phylogeny, soils and climate. Biogeosciences 6, 2677-2708.

Phillips OL, van der Heijden G, Lewis SL, López-González G, et al (2010) Drought-mortality relationships for tropical forests. New Phytologist 187:631-646.


Fyllas NM, Dimitrakopoulos PG, Troumbis AY, 2008. Regeneration dynamics of a mixed Mediterranean pine forest in the absence of fire. Forest Ecology and Management 256, 1552-1559.


Fyllas NM, Phillips OL, Kunin WE, Matsinos YG, Troumbis AI, 2007. Development and parameterization of a general forest gap dynamics simulator for the North-eastern Mediterranean Basin (GREek FOrest Species). Ecological Modelling 204, 439-456.